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The Indian leopard ( Panthera pardus fusca) is a of the ( P. pardus). It is widely distributed on the Indian subcontinent. It is threatened by of skins and body parts, and persecution due to human-leopard conflict and retaliation for depredation.


Taxonomy
Felis fusca was the proposed by Friedrich Albrecht Anton Meyer in 1794 who described a black leopard from that was on display at the Tower of London. Leopardus perniger proposed by Brian Houghton Hodgson in 1863 were five leopard skins from , out of which three were . He mentioned and Nepal as habitat. Panthera pardus millardi proposed by Reginald Innes Pocock in 1930 was a single leopard skin and skull from . It differed from typical P. p. fusca skins by longer hair and a more greyish colour.

Since leopard populations in Nepal, Sikkim and Kashmir are not geographically isolated from leopard populations in the Indian subcontinent, they were subsumed to P. p. fusca in 1996. The in the west and the in the north form barriers to the dispersal of this subspecies. In the east, the and the lower course of the Brahmaputra River are thought to form natural barriers to the range of the Indochinese leopard.

A of 49 leopard skin samples collected in Azad Jammu Kashmir and regions of northern Pakistan revealed of both Persian and Indian leopards, indicating of both subspecies in this region.


Characteristics
The Indian leopard has strong legs and a long, well-formed tail, broad muzzle, short ears, small, yellowish-grey eyes, and light-grey ocular bulbs. Its coat is spotted and rosetted on a pale yellow to yellowish-brown or golden background, except for the forms; the spots fade toward the white underbelly and the insides and lower parts of the legs. Rosettes are most prominent on the back, flanks and hindquarters. The pattern of the rosettes is unique to each individual.
(2025). 9789350097618, Hachette India.
Juveniles have woolly fur, and appear dark due to the densely arranged spots. The white-tipped tail is long, white underneath, and displays rosettes, which form incomplete bands toward the end. The rosettes are larger in other leopard subspecies in Asia. Fur colour tends to be more pale and cream in arid habitats, more grey in colder climates, and of a darker golden hue in habitats.

The clouded leopard can be told apart by its diffuse "clouds" of spots compared to the smaller and distinct rosettes of the leopard, longer legs and thinner tail.

(2025). 9781845377359, New Holland.


Skull
The largest skull for an Indian leopard was recorded in 1920, and belonged to a large, cat in the area of , . The black panther was said to have bigger forelimbs and forequarters than hind-limbs and hind-quarters, with a skull and claws nearly as large as those of a tigress. The skull measured in basal length, and in breadth, and weighed . By comparison, the skull of one western measured in basal length, and in breadth, and weighed .


Size
Male Indian leopards grow to between and in body size with a to long tail and weigh between . Females are smaller, growing to between and in body size with a to long tail, and weigh between . Sexually dimorphic, males are larger and heavier than females.

The largest wild individual appears to have been a male man-eater that was shot in the Dhadhol area of Bilaspur district, Himachal Pradesh in 2016. It reportedly measured from head to tail, at the shoulder, and weighed .


Distribution and habitat
The Indian leopard is distributed in , , and parts of . Bangladesh has no viable leopard population but there are occasional sightings in the forests of , Chittagong Hill Tracts and Cox's Bazar. It inhabits tropical rainforests, dry , and northern coniferous forests but does not occur in the of the .

In southern , it was recorded in Qomolangma National Nature Preserve.

In Nepal's Kanchenjunga Conservation Area, a melanistic leopard was photographed at an elevation of by a camera trap in May 2012.


Population in India
In 2015, 7,910 leopards were estimated to live in and around tiger habitat in India; about 12,000 to 14,000 leopards were speculated to live in the entire country. The following table gives the major leopard populations in the Indian states. As of 2020, the leopard population within forested habitats in India's tiger range landscapes was estimated at 12,172 to 13,535 individuals. Surveyed landscapes included elevations below in the and , and , , as well as the Brahmaputra River basin and hills in . As per 2022, the Indian leopard population was estimated at 13,874 individuals.

+Leopard population by state
569
42
74
86
722
77
51
1,879
570
3,907
1,985
568
721
1070
297
371
652
233
13,874


Behaviour and ecology
The leopard is elusive, solitary, and largely . It is known for its ability in climbing, and has been observed resting on tree branches during the day, dragging its kills up trees and hanging them there, and descending from trees headfirst.Jerdon, T.C. (1874). Mammals of India: a natural history of the animals known to inhabit continental India. John Wheldon, London. It is a powerful swimmer, although is not as disposed to swimming as the tiger. It is very agile, and can run at over , leap over horizontally, and jump up to vertically. It produces a number of vocalizations, including grunts, roars, growls, meows, and purrs.
(1991). 9780520080850, The University of California Press. .

In Nepal's Bardia National Park, home ranges of male leopards comprised about , and of females about ; female home ranges decreased to when they had young cubs. In Gir National Park, the home range of a male radio-collared leopard was estimated at . It killed prey once in 3.7 days.

The leopard is a versatile, opportunistic hunter, and has a very broad diet. It is able to take large prey due to its massive skull and powerful jaw muscles.

(2025). 9780789477644, DK Adult.
(1984). 9780671428051, Simon & Schuster/Touchstone Books. .
In Sariska Tiger Reserve, the dietary spectrum of the Indian leopard includes , , , , , and . In Periyar Tiger Reserve, make up a large proportion of its diet.


Reproduction
Depending on the region, the leopard mates all year round. The lasts about 46 days and the female usually is in heat for 6–7 days. Gestation lasts for 90 to 105 days. Cubs are usually born in a litter of 2–4 cubs. Mortality of cubs is estimated at 41–50% during the first year. Females give birth in a cave, crevice among boulders, hollow tree, or thicket to make a den. Cubs are born with closed eyes, which open four to nine days after birth.
(2025). 9780226779997, University of Chicago Press.
The fur of the young tends to be longer and thicker than that of adults. Their pelage is also more grey in colour with less defined spots. Around three months of age, the young begin to follow the mother on hunts. At one year of age, leopard young can probably fend for themselves, but remain with the mother for 18–24 months. The average typical life span of a leopard is between 12 and 17 years.


Sympatric carnivores
Indian leopards are not common in habitats where tiger density is high, and are wedged between prime tiger habitat on the one side, and cultivated village land on the other. Where the tiger population is high or increasing, tigers drive leopards off to areas located closer to human settlements, like in Nepal's Bardia National Park and Sariska Tiger Reserve. Resource partitioning occurs where leopards share their range with tigers. Leopards tend to take smaller prey, usually less than , where tigers are present. In areas where leopard and tiger are sympatric, coexistence is reportedly not the general rule, with leopards being few where tigers are numerous. The mean leopard population density decreased significantly from 9.76 to 2.07 animals per , while the mean tiger population density increased from 3.31 to 5.81 animals/100 km2 from 2004–2005 to 2008 in Rajaji National Park following the relocation of pastoralists out of the park. There, the two species have high dietary overlap, and an increase in the tiger population resulted in a sharp decrease in the leopard population and a shift in the leopard diet to small prey (from 9% to 36%) and domestic prey from 6.8% to 31.8%. In Chitwan National Park, leopards killed prey ranging from less than in weight with most kills in the range. Tigers killed more prey in the range. There were also differences in the microhabitat preferences of the individual tiger and leopard followed over five months; the tiger used roads and forested areas more frequently, while the leopard used recently burned areas and open areas more frequently. When a tiger killed baits at sites formerly frequented by leopards, the leopards did not hunt there for some time.

In the tropical forests of India's Nagarhole National Park, tigers selected prey weighing more than , whereas leopards selected prey in the range. In tropical forests, they do not always avoid the larger cats by hunting at different times. With relatively abundant prey and differences in the size of prey selected, tigers and leopards seem to successfully coexist without competitive exclusion or interspecies dominance hierarchies that may be more common to the leopard's co-existence with the lion in savanna habitats. In areas with high tiger populations, such as in the central parts of India's Kanha National Park, leopards are not permanent residents, but transients. They were common near villages at the periphery of the park and outside the park. In a reserved forest of southern India, species preyed upon by leopard, and overlapped considerably.

The leopard and both hunt and , but the leopard usually prefers forested habitats located at lower elevations. Leopard may conflict with and can follow them up trees. Bear cubs are probably far more vulnerable and healthy adult bears may be avoided by leopards. One leopard killed a three-quarters grown female sloth bear in an apparently lengthy fight that culminated in the trees. Apparently, a sloth bear killed a leopard in a confrontation in Yala National Park, but was itself badly injured in the fight and was subsequently put down by park rangers.Kurt, F., & Jayasuriya, A. (1968). Notes on a dead bear. Loris, 11, 182–183.


Threats
Hunting of Indian leopards for the illegal is the biggest threat to their survival. They are also threatened by loss of habitat and fragmentation of formerly connected populations, and various levels of human–leopard conflict in human–dominated landscapes.

Several newspapers reported of leopards falling into open wells and being rescued with the help of Forest Department officials.


Poaching
A significant immediate threat to wild leopard populations is the illegal trade in poached skins and body parts between India, Nepal and . The governments of these countries have failed to implement adequate enforcement response, and wildlife crime remained a low priority in terms of political commitment and investment for years. There are well-organised gangs of professional poachers, who move from place to place and set up camp in vulnerable areas. Skins are rough-cured in the field and handed over to dealers, who send them for further treatment to Indian tanning centres. Buyers choose the skins from dealers or tanneries and smuggle them through a complex interlinking network to markets outside India, mainly in China.Banks, D., Lawson, S., Wright, B. (eds.) (2006). Skinning the Cat: Crime and Politics of the Big Cat Skin Trade . Environmental Investigation Agency, Wildlife Protection Society of India Seized skins in confirm the city's role as a key staging point for illegal skins smuggled from India bound for Tibet and China.Banks, D. (2004). The Tiger Skin Trail. Environmental Investigation Agency.

It is likely that seizures represent a tiny fraction of the total illegal trade, with the majority of smuggled skins reaching their intended end market. Seizures revealed:

  • in China and Tibet: nearly 130 leopards were killed every year, e.g. more than 774 poached leopards between July 1999 and September 2005.


Human–leopard conflict

Causes of conflict
Expansion of used land, encroachment by humans and their livestock into protected areas are main factors contributing to habitat loss and decrease of wild prey. As a result, leopards approach human settlements, where they are tempted to prey on dogs, pigs and goats – domestic livestock, which constitutes an important part of their diet, if they live on the periphery of human habitations. Human–leopard conflict situations ensue, and have increased in recent years. In retaliation for attacks on livestock, leopards are shot, poisoned and trapped in snares. The leopards are considered to be unwanted trespassers by villagers. Conservationists criticize these actions, claiming that people are encroaching on the leopard's native habitat.Sears, S. (2008). "Mumbai Leopards: Killers or Victims?" Wildlife Extra, 11 April 2008.Sears, S. (2009). "The wild leopards of Oman and Nepal – And how to see them" . Wildlife Extra, April 2009. India's Forest Department is entitled to set up traps only in cases of a leopard having attacked humans. If only the presence of a crowd of people prevents the leopard from escaping, then the crowd has to be dispersed and the animal allowed to escape.Athreya, V., Belsare, A. (2007). Human – Leopard Conflict Management Guidelines. Kaati Trust, Pune, India.

As urban areas expanded, the natural habitats of leopards shrunk resulting in leopards venturing into urbanized areas due to easy access of domestic food sources. Karnataka has a high number of such conflicts. In recent years, leopards were sighted in , and the forest department captured six leopards in the city's outskirts, relocated four of them to various other locations.


Man-eater leopards
Every year more leopards are killed by humans than the humans killed by leopards. On average nearly 400 leopards are reported killed yearly in India, Nepal and China combined based on the leopard skins caught from the poachers, though the actual number of leopards killed by humans is likely to be several times higher. In and around the of alone, 68 leopards were killed by people between 2001 and 2013, of which only 10 were man-eaters.

The frequency of Leopard attacks on humans varies by geographical region and historical period. Since India and have the majority of Indian leopards population, consequently attacks are regularly reported only from India and Nepal. Among the five "", leopards are less likely to become man-eaters—only and have a less fearsome reputation.

(2025). 9780521825054, Cambridge University Press. .
While leopards generally avoid humans, they tolerate proximity to humans better than and and often come into conflict with humans when raiding livestock.
(2025). 9780393326093, W. W. Norton & Company. .

Attacks in India are still reported, since leopards population in India outnumber population of all other large carnivores combined, consequently the number of humans killed by leopards is also more than those killed by all other large carnivores combined.

In Nepal, where most attacks occur in the midland regions, i.e. in the , midhills, and , the rate of leopard predation on humans results in approximately 1.9 human deaths annually per million inhabitants. .

Historically, with rapid urbanization in late 19th and early 20th centuries, leopard attacks may have peaked in India during those times. Notable man-eaters of that era include Leopard of Central Provinces, Rudraprayag, Gummalapur, Yellagiri Hills, Golis Range and Panar.


Ways to minimise conflict
Key to avoiding conflict or leopard's predation of humans is to shift the focus on human's behavioral change to minimise the chances of a leopard encounter or attack, which can be achieved by "clearing bushes and overgrowth around homes to minimise hiding spaces for leopards, leaving a light on at night to deter them, and ensuring people, especially children, did not go out alone at night." Leopards are shy and avoid humans and are more active at night, during encounter with leopards "give way to the leopard and move away calmly" and alert the forest department immediately. Spotted in India: Humans and leopards living in harmony , ourbetterworld.org, 9 Apr 2021.


Conservation
Panthera pardus is listed in CITES Appendix I. Despite India and Nepal being contracting parties to , national legislation of both countries does not incorporate and address the spirit and concerns of CITES. Trained human resources, basic facilities and effective networks for control of poaching and trade in wildlife are lacking. The Indian leopard is considered Vulnerable in India, Bhutan,
(2025). 9789998046023, National Environment Commission, Royal Government of Bhutan. .
and Nepal
(2025). 9780900881602, Department of National Parks and Wildlife Conservation. .
but Critically Endangered in Pakistan.

Frederick Walter Champion was one of the first in India who after World War I advocated for the conservation of leopards, condemned sport hunting and recognised their key role in the ecosystem.

(2025). 9788185019536, Natraj Publishers.
Billy Arjan Singh championed their cause since the early 1970s.
(1982). 9780195654028, Jonathan Cape.

There are a few leopard rescue centres in India, such as the Manikdoh Leopard Rescue Centre in , but more rescue and rehabilitation centres are being planned. Some wildlife experts think that such centres are not an ideal solution, but that conflict resolution by way of changing human behaviour, land use or grazing patterns and implementing responsible forest management to lessen human-animal conflict would be far more effective to conserve leopards.


In culture and literature
  • An Indian named '' is featured in 's 1894 novel The Jungle Book, as well as in 's 1967 and 2016 film adaptations.
  • The book Man-Eaters of Kumaon is based on man-eating leopards and tigers in .
  • is a 2014 directed by and written by Gauri Bapat. It is supposedly based on true events.
  • Local people in the call a darkish grey panther pogeyan, which purportedly lives in the high-elevation grasslands of the Anamudi forest.


See also


Further reading

External links

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